Hemizygotes arrest development prior to the third instar larval stage.

Oogenesis is completed and eggs are laid in females containing homozygous germline clones, however, the embryos fail to hatch. The mutant embryos show severe disorganisation of both the DNA and microtubules and no structures identifiable as pronuclear fusion or syncytial divisions are observed. Chromosomes are dispersed throughout the cytoplasm and microtubules form large asters surrounding the dispersed chromosomes.

Late-stage oocytes show defects in mutant females. Firstly, spindle microtubule bundles are much longer than in wild type. Secondly, ooplasmic microtubules are more numerous and longer than in wild type and are often arranged in a starburst pattern. Spindle disorganisation ranging from slight (bipolar with some fraying or bending) to extreme (long microtubule bundles with no apparent poles) is seen. The central spindle region is often missing in the mutant oocytes, resulting in two disconnected half spindles and multipolar spindles. The karyosome and spindle are not in proper contact. Bi-orientation of homologous chromosomes is defective.