The morphology of the Golgi stacks is severely disrupted in mutant spermatocytes, with the Golgi consisting of clusters of irregularly sized and swollen cisternae. Mutant onion-stage spermatids lack acroblasts and display numerous vesicles scattered throughout the cytoplasm. The structure of the basal body and centriolar adjunct is normal in mutant onion-stage spermatids, but in 87% of cases they are displaced from the nuclear envelope. In mutant spermatids, 25% of mutant axonemes appear irregular in cross-section and lack peripheral doublets and/or central microtubules.

Mild defects in cytokinesis are seen in neuroblasts in mutant larval brains: 15% of mid-telophases have poorly constricted or broken contractile rings. The mitotic index and frequency of prometaphases/metaphases relative to all mitotic figures are not significantly different from wild type.