Frameshift deletion (23bp) of 4:480801 through 480823 resulting in the addition of SSS after isoleucine 281 of Asator PD, PE, and PG isoforms (which corresponds to isoleucine 280 of the PI and PJ isoforms and isoleucine 125 of the PF, PH, PK, and PL isoforms) and premature truncation after these novel amino acids.