FB2026_03 , released September 17, 2026
CV Term Report
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Term transmedullary neuron Tm2 ID (Ontology) FBbt:00003790 (Fly Anatomy)
Definition Transmedullary narrow field neuron that has its main dendritic arborization in medulla layer M2 (Ting et al., 2014), where it receives input from lamina monopolar neuron L2 (Takemura et al., 2011), and additional postsynaptic terminals in M4-M5, where it receives input from lamina monopolar neuron L4 (Takemura et al., 2011). It has presynaptic terminals in layers M3, M4 and M9 (Fischbach and Dittrich, 1989; Takemura et al., 2011). Its axon follows the posterior edge of its cognate medulla column (Ting et al., 2014), then crosses the second optic chiasma to terminate in lobula layers 1 and 2 (Fischbach and Dittrich, 1989). In layer 1 it outputs to T5 neurons (Shinomiya et al., 2014). It is a cholinergic neuron (Varija Raghu et al., 2011; Gao et al., 2008; Takemura et al., 2011). There is usually one of these cells per optic column (Nern et al., 2025).[ FlyBase:FBrf0049410 FlyBase:FBrf0204652 FlyBase:FBrf0206213 FlyBase:FBrf0212356 FlyBase:FBrf0217052 FlyBase:FBrf0222324 FlyBase:FBrf0224196 FlyBase:FBrf0225094 FlyBase:FBrf0262545 ]
Also Known As"Tm2"
CommentThe neurotransmitter has been assessed by co-labelling cells with a P\{Cha-GAL4.7.4\} driver (FBtp0014830) and an anti-ChAT antibody (Varija Raghu et al., 2011), by using a Cha-GAL4.DBD (FBtp002350) (Gao et al., 2008) and by neurotransmitter was assessed by single cell RT-PCR measurement of choline acetyltransferase (FBgn0000303) expression (Takemura et al., 2011). Connectivity in the medulla was extrapolated from serial EM of 3 columns from a single sample (Takemura et al., 2008) or from 7 columns (Takemura et al., 2013). Connectivity in the lobula was assessed by serial section electron microscopy of a single column (Shinomiya et al., 2014).
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VFB:FBbt_00003790
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Insertions (FBti)EXPRESSION_STAGE_TISSUE_POSITION
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 transmedullary neuron Tm2 (all annotations which use CV term, excluding "NOT" statements)
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transmedullary neuron
 |__narrow field transmedullary neuron_____
cholinergic neuron                         |
 |__adult cholinergic neuron_______________|
adult neuron                               |
 |__adult cholinergic neuron_______________|
secondary neuron                           |
 |__Notch ON hemilineage secondary neuron__|
Notch ON hemilineage neuron                |
 |__Notch ON hemilineage secondary neuron__|
optic lobe narrow field columnar neuron    |
 |__narrow field transmedullary neuron_____|
                                           transmedullary neuron Tm2  36 rec.
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Is a narrow field transmedullary neuron
Notch ON hemilineage secondary neuron
adult cholinergic neuron
Develops from immature transmedullary neuron Tm2
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FlyBrain_NDB:10144
VFB:FBbt_00003790