In aret^PA/aret^WH adult females, oogenesis proceeds normally until stage 4 - meiosis and mitosis of germ cells occurs normally and nurse cells become enlarged an polyploid. But during stage 4, when mitosis of germ cells is over in wild-type, in around 20% of cysts the nurse cells condense their chromosomes and form large mitotic spindles.

Egg chambers from aret^WH/aret^WH females contain far greater than the normal 16 germ cells, and these germ cell fail to become polyploid. Marker analysis suggests that these germ cell fail to exit mitosis at stage wild-type germ cells do.

Numerous necrotic cells are seen in mutant ovariole at high frequency. Germline cells in mutant ovarioles undergo complete cytokinesis, but never form ring canals. No fusome is present in these cells, tiny and mispositioned spectrosome remnants are seen. Germline development in mutants pre-oogenesis is wild-type.

No differentiating egg chambers.