Hemizygous and homozygous germline clone cells are viable.

Semilethal, survivors show a rough reduced dark eye colour and lack a variable number of orbital, ocellar and vertical bristles, vibrissae are sparse, wings have thickened veins and incisions along inner margins.

Semilethality is polyphasic, occurs during larval, pupal and adult stages. Second and third instar larvae grow slowly, those that survive to pupariate and hatch imagos. Mosaic analysis demonstrates mutant phenotype is non-autonomous. Individuals survive when Dp(1;4)w^m65g, Dp(1;3)w^m49a or Dp(1;3)N^264-58 is present. Surviving males have rough, reduced darkened eyes, vibrissae are sparse and orbital, ocellar and vertical bristles often absent. Wing veins are thick and the inner wing margin has incisions. Males are sterile.

Eye colour: dark in adult escapers.