Embryos derived from grk²/grk⁶ females gastrulate with two distinct ventral furrows. Cuticles show an expansion of ventral structures.

grk¹/grk² heterozygous females display fused egg chambers and gaps in the follicular epithelium that uncover the nurse cells. Eggs derived from these females exhibit strong ventralised phenotype.

Ventralising mutation, eggs are longer and less rounded than wild type and a single dorsal eggshell filament is formed on the dorsal midline. Egg chambers from grk²; bwk⁰⁸⁴⁸² females also have single dorsal filaments but they resemble bwk filaments, short and ragged.

grk^2B6/grk² heterozygous eggs lack dorsal ventral polarity and possess a posterior micropyle. The eggs never contain embryos.

Intermediate and strong ventralised embryos. orb^F343 and orb^dec interact genetically with grk, double heterozygote embryos display fused dorsal appendages.

Follicular epithelium of mutant egg chambers has holes.

Reduction or absence of dorsal appendages (ventralization) with increase in main body egg shell. Micropyle at both ends of eggs, embryos ventralized.

An enlarged mass of mesodermal cells invaginates on the ventral side of the embryo, the enlarged mesoderm is often organised into two ventral furrows. Organisation is lost in later stages of development and a mass of mesodermal cells fills the ventral half of the embryo. Respiratory appendages are fused and inserted directly on the dorsal midline.

phenotype of eggs and embryos: weak and intermediate