Homozygous tracheal terminal cells show a reduction in branching compared to wild type. The mutant cells appear to have normal gas-filled lumens.

When somatic clones are made in the follicular epithelium cell polarity defects are seen. These are more severe at the poles of the egg chamber than in lateral positions where polarity is often normal. Zonula adherens is also compromised in clones. Mutant follicle cells are also multilayered. most strikingly at the anterior and posterior poles. Border cell clusters that contain both normal and mutant cells migrate at the correct stages, but there are frequent abnormal arrangements of rosette and central polar cells. Border cells are often elongated with individual cells trailing each other rather than migrating as a group of closely attached cells. Those cells that trailed behind often have elongated extensions which contact anterior cells.

Male embryos are convoluted and are not dorsally closed.