TDT attachment sites vary. The muscle may be found in the wild type position, posterior to the intrascutal suture and displaced medially, anterior to the intrascutal suture or missing entirely. Muscle may also be reduced in size or split dorsally attaching to two separate sites on the scutum. Cytology of the TDT is also altered, muscles may have fibres that are swollen and stain abnormally, other fibres may have large, axially aligned holes. Attachment pattern of the DVMs is also altered. DVMs remain within their respective attachment regions or extend into the region normally occupied by an adjacent DVM. DVMs may be entirely missing. Many flies fail to eclose, dying during late pupal stages. Flies that do eclose show impaired motility. Flies can get stuck in their food, falling into the food causes their femurs to become swollen and bent.