Homozygous adults have abnormal bristles. Onion-stage spermatids usually contain a single Nebenkern of larger than normal size, associated with two or more normal-sized nuclei (25.3% of spermatids have 2 nuclei, 49.2% have 4 nuclei). Onion-stage spermatids of chic³⁷/chic^R1 males usually contain a single Nebenkern of larger than normal size, associated with two or more normal-sized nuclei (67.6% of spermatids have 4 nuclei). Onion-stage spermatids of chic⁰¹³²⁰/chic^R1 males usually contain a single Nebenkern of larger than normal size, associated with two or more normal-sized nuclei (64.3% of spermatids have 4 nuclei). The central spindle is severely defective in the first meiotic division in homozygous males, and the actin-based contractile ring is missing. Abnormal aggregations of F-actin are seen in premeiotic mature spermatocytes at the S5 stage. These aggregates may represent remnants of the fusome. 63.9% of asters show irregular positioning and delayed migration during meiotic divisions in homozygous males. 87% of asters show irregular positioning and delayed migration during meiotic divisions in chic³⁷/chic^R1 males. 69.5% of asters show irregular positioning and delayed migration during meiotic divisions in chic⁰¹³²⁰/chic^R1 males.