Nuclei are often mispositioned in cuprophilic cells in the midgut of mutant third instar larvae, with 13-71% of the nuclei in each gut being found at the outer edges of the cuprophilic cells, causing a bulge near the pore. Pore size is normal in the mutant cuprophilic cells.

The gut lumen is acidified in mutant first instar larvae. However, mutant third instar larvae show a statistically significant decrease in overall gut acidification compared to controls.

Mutant wing cells produce normal-looking hairs, but the hairs are often found on a small pedestal. The wing cell surface that is not hair is rough and remnants of cell outlines are sometimes visible.

The border cells migrate ahead of the follicle cell monolayer in 73% of mutant egg chambers, in contrast to wild-type where the border cells and follicle cell monolayer migrate in a concerted fashion. Egg chambers where the border cells are retarded relative to the follicle cells are occasionally seen. Most mutant oocytes occupy a larger portion of the egg chamber than in wild type during follicle cell migration, while the oocyte shows no significant overgrowth at the completion of migration. Most mutant oocytes occupy a larger portion of the egg chamber than in wild type during border cell migration. Some follicle cells often remain in contact with the nurse cell membranes at stage 10A. These follicle cells still attempt to make the appropriate adhesive contacts with the oocyte membrane, pulling the oocyte membrane towards them and grossly distorting the nurse cells/oocyte interface. In most cases, the subsequent inward migration of the follicle cell layer at stage 10B proceeds relatively normally, but in rare, extreme cases, the centripetally migrating cells penetrate between nearby nurse cell membranes and cause one or more nurse cells to become included within the egg along with the oocyte. Mutant follicle cells often appear more cuboidal than wild-type follicle cells. There is no overproliferation in the follicle monolayer. The majority of follicle cells fail to constrict their apices over the oocyte as they migrate, in contrast to wild type. The mean apical surface area of mutant follicle cells is almost twice that of wild type. Comparison of the apical surface areas of mutant follicle cells in chambers during migration with those where migration has been completed reveals a slight increase, suggesting that the monolayer cannot withstand the forces exerted by the growing oocyte (in addition to the constriction defect). In approximately 10% of egg chambers, migratory cells that are well separated from, or trail behind, the main border cell cluster are seen.