Under standard culture conditions, few homozygous adults are recovered. If hemizygous male larvae are cultured under uncrowded conditions, 60-80% develop into adults, but grow slowly (the mutants take approximately 3 days longer to develop compared to the normal developmental time). The mutant males have 35% reduced weight compared to controls. Mutant wing cells show a 13% reduction cell width compared to controls. The macrochaetae are thinner and shorter than normal.

Homozygous clones in the imaginal discs consistently contain fewer cells than their wild-type sister clones.

Homozygous females lay eggs, but the eggs do not hatch. The embryos derived from homozygous females show irregular distribution of cleavage nuclei. Embryos are often arrested around cycle 8, and most of the nuclei are at M-phase. Some polyploid nuclei are seen. Some embryos continue development past gastrulation.

The mitotic index in the larval brain of hemizygous males is reduced compared to controls.