In late third instar larva, strong expression in the lamina anlagen, outer optic anlagen, in neuroblasts of the central brain; patches of expression in wing, haltere, and leg discs.
Larval expression is observed in subset of Kenyon cells in each of four clusters, in the calyx and pedunculus in a two-fold concentric organization, and in the cap and ring of the α and α' β and γ lobes of the mushroom body.