Subject: Dp(4;3)f Personal communication from: Kathy Matthews, Bloomington Drosophila Stock Center Subject: Dp(4;3)f Information communicated: The aberration described as Tp(4;3)f is really Dp(4;3)f. The aberration was originally described as a T(3;4) and was apparently renamed as a transposition based on the cytology reported by Lewis (1956, DIS 30:130 ): 'Insertion of at least seven bands of chromosome four (bands not identified) into 3L, probably just after 65D1-2.' Lewis did not, however, describe a reciprocal 4th chromosome deficiency. The two extant stocks (Bloomington numbers 894 & 895) came to us as Tp(4;3)f with a 3rd chromosome balancer specified but no 4th chromosome homologue specified. I tested stock number 895 for the presence of the duplication and deficiency segregants and found only the duplication. The incorrect genotypic information has led to some confusion so I'll summarize the complementation data for this duplication below. The genotypes tested by Rui Sousa-Neves (see FBrf0112252 & FBrf0125105) and me were as follows: Dp(4;3)f/+; svspa-1 The duplication complements svspa-1 (this genotype has a wild- type eye); Sousa-Neves (FBrf0112252) & Matthews (this pc) Dp(4;3)f/+; T(1;4)scH/pho1 The duplication fails to complement the lethality of T(1;4)scH/ pho1 (this genotype is lethal); Sousa-Neves (FBrf0125105; the genotype tested was clarified in discussion with me) Dp(4;3)f/+; In(4)ciD, ciD panciD/pan3 The duplication fails to complement panciD/pan3 (this genotype is lethal); Sousa-Neves (FBrf0112252) Thus, Dp(4;3)f is sv+, pho- and pan-. It is not known whether it is ci+ or ci-. Tp(4;3)f and Df(4;3)f almost certainly do not exist.