Mutant pupae show defects in leg disc morphogenesis. The T1 and T2 legs appear at the proper times relative to wild-type. However mutants reveal initial mislocalisation of one T1 leg, which is only seen in a small number of wild-type pupae. Mutant legs also fail to form the clearly ordered array that is evident on completion of eversion in wild-type prepupae and they undergo no significant elongation after 8 hours after puparium formation. As a result the final leg elongation that occurs at head eversion, which is delayed in mutants to between 15:30 and 26:00 hrs after puparium formation, results in legs that are significantly shorter than those seen in wild-type pupae. The legs are malformed at this stage, with one of the third legs wrapped around the edge of the wing.

2-3% of crol⁰⁶⁴⁷⁰/crol^k08217 mutant animals are able to eclose from the pupal case, though these adults have severe morphological defects. 71% of these escapers have strongly misshapen legs, often with a severe kink in the femur. All escapers also display wing defects. Over half have held out wings with a partial (blister) or complete (balloon) separation of the dorsal and ventral wing surfaces, while the remainder have either malformed or completely unfolded wings. The blisters seen are generally large and do not appear to have sharp boundaries.

Homozygotes display a lethal phenotype indistinguishable from that of crol⁰⁴⁴¹⁸.

crol^k08217 is an enhancer of wing phenotype of mys⁸

crol^k08217 is an enhancer of wing phenotype of if³