Eggs laid by grk⁶/Lis-1^3.1.2 mothers show defects in the location and morphology of the dorsal appendages. Most dorsal appendages are fused. The slight loss of viability phenotype seen in Lis-1^k11702/Lis-1^3.1.2, is enhanced by the addition of Gl¹. The adult escapers from this cross generally do not survive more than a few days. The addition of this Lis-1 transheterozygote combinations enhances the eye phenotype seen in Gl¹ flies. In addition scutellar bristles of escapers flies are reduced in size and frequently lost. Most adult escapers lack bristles on the ventral side of the abdomen, and in many animals abdominal tergites are completely absent in one or more segments.