Viable in transheterozygous combination with Hsp839J1; males are sterile and females are weakly fertile. Hsp839J1/Hsp83e4A males show some defects during spermatogenesis. The number and shape of spermatocytes within 16-cell cysts are mostly normal (5-10% are abnormal). The nuclei and nebenkern within spermatids are mostly normal (5-10% are abnormal). Individualised sperm are present, but are not motile. Hsp8308445/Hsp83e4A males show defects during spermatogenesis. Excessive numbers of primary spermatocytes are seen in many developing cysts. Spermatids with variable number, size and shape of nuclei and nebenkern are seen. Individualised sperm are present but they are not motile and are fragile.
6+/-2% of KrIf-1/+ flies born to Hsp83e4A/+ mothers have outgrowths with ectopic vibrissae protruding from the ventral region of the eye, compared to less than 0.1% of KrIf-1/+ flies born to isogenised wild-type mothers.